From Seville: a full day
The listing gives about 10 hours, with pickup in central Seville, transport to Doñana and a stretch by 4x4 or minibus through the northern area of the park. The itinerary adds birdwatching at Dehesa de Abajo, the village of El Rocío, a walking route, time for lunch and the dunes of the Huelva coast. The departure time changes with sunrise and the local provider confirms it after booking.
From El Rocío: the park only
The 4x4 tour leaves from El Rocío and takes between three and four hours according to the listing. It covers the northern area, including Coto del Rey, with a stop at the José Antonio Valverde Visitor Centre. It is the short option if you are already in El Rocío or moving around the area on your own.
What the park administration says
Spain's Ministry for the Ecological Transition says the park interior is covered by three itineraries, two of them overland in off-road vehicles, run by concessionaire or authorised companies with guides. For the northern itinerary, between Coto del Rey and the José A. Valverde Visitor Centre, its official page says six authorised companies run it and that you must book in advance.
Frequently asked questions
Seville or El Rocío: which one should I pick?
If you are staying in Seville and want a full day with birds, the village and the dunes, take the departure from Seville. If you only want the park by 4x4 and are near El Rocío, take the tour of between three and four hours.
Is cancellation free?
The Seville listing gives free cancellation up to 24 hours before; the El Rocío listing gives free cancellation up to 48 hours before.